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Dated May 15 2017 BIOTECHNOLOGY VALUE FUND L.P. BVF INC. By BVF Inc. its general partner President MARK N. LAMPERT BVF PARTNERS L.P. CO BVF Partners L.P. 3 266 321 PN IA BVF Inc. Mark N. Lampert United States IN Item 1 a. Address of Issuer s Principal Executive Offices 280 East Grand Avenue South San Francisco California 94080 Item 2 b. Ownership of More than Five Percent on Behalf of Another Person. Partners BVF Inc. and Mr. Lampert share voting and dispositive power over the shares of...

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